Only one large scale paper has actually been written about Fukuisaurus. Perhaps the minimal material, though there is some more material now than there was when initially described, has led to the lack of published research concerning Fukuisaurus. That paper, unfortunately, is published in the Journal of Vertebrate Paleontology; meaning that without a subscription it is virtually unreadable in its entirety. If you have a subscription or would like to buy the article it can be found here. If you do not have a subscription or wish to buy the article, you do not have to, of course. Remember that the description of the genus was conducted with only two bones of the skull; a right maxilla and jugal. Since that time, when it was described as a "non-hadrosaurid iguanodontian", more skull material has been collected and the lack of kinetic movement in the skull through fusion and other characters in a cladistic analysis (no published version of this information appears to be available) showed it to actually be a basal hadrosaur. The fact that this information is not known to be published means it may be someone's research that is still in progress or an as yet peer reviewed study that has assigned it. Hopefully a reference to this assertion will be found at some point and it will not continue on as hearsay one way or another.
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Fukuisaurus Described
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Fukuisaurus and its Jaw
It is strange how little of Fukuisaurus has been recovered. It is also strange how pictures of the bones are not in circulation; this is no doubt in part due to the exclusivity of JVP articles and the images used therein.The jaw fragments used in the description are present in the model, as casts of course, on exhibit in Fukui Prefecture's museum and we can therefore see them there. They present as expected; as typical hadrosaurine or iguanodontid jaw bones would be expected. Really, the descriptive material is minimal, but consistent with the types of dinosaurs that Fukuisaurus is reported to be related to. The extrapolations, thumb spikes, hadrosaurine body head and posture, are within reason given the diagnosis and description; however, what really needs to be done is more excavation. The only thing that will truly cement the placement and validity of Fukuisaurus is more evidence that attests to the certainty of the current taxonomic placement of the animal. Ideally an entire skeleton may be found, but for now I think that the cause for the skeleton would be furthered by even smaller associated numbers of material such as an entire skull or bits of forelimbs and/or hindlimbs discovered near known skull elements. More expeditions, digging, and a little luck will go a long way and can only be seen in time though.
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Model near the skeletal model in Fukui Prefecture's museum |
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Fukuisaurus for Everyone
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The wary kind are more dangerous it seems. |
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More Mimics
Photo by Scott Anselmo |
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Comparing the Family
Scott Hartman does wonderful things and today, this image, is exactly what we are looking for. Not only does it present the known remains of Suchomimus, our dinosaur in question this week, but it also shows the known evidence of its extended family. The skulls alone are very good examples of why these three animals have been placed in the same lineage. "Lumpers", those in favor of placing fossil specimens in as few species as possible through comparisons between currently separated species, could use these sets of remains to make a rather strong argument for a growth series, given only the amount and types of bones that exist and comparing them to one another. The major difference between the three that would be most difficult to reconcile would of course be the dorsal neural spines of Spinosaurus. The skulls are all similarly shaped, however, and the hands and forelimbs that we have for Baryonyx as well as Suchomimus are very similar. Hindlimb evidence is missing for Spinosaurus and Baryonyx, though we have a very little amount of Baryonyx material, whereas it is quite well documented in Suchomimus; pelvic elements of Suchomimus and Baryonyx are well represented. The majority of the vertebrae are absent in all three species and ribs and gastralia are only well evidenced in Suchomimus though some do exist in association with Baryonyx remains.
The first thing anyone notices with regard to Spinosaurs like Baryonyx and Subhomimus is the posture. The reason that people always notice the posture is because it is always the same in mounted specimens, casts, and models. Hindlimbs are slightly bent, tail straight out, torso angled downward and forward, with the mouth agape and the forelimbs splayed open, palms of the hands facing one another. The constantly used posture leads to confusion over the type of animal on display. Baryonyx being more widely known than Suchomimus, many Suchomimus skeletons are misnamed and mistaken as Baryonyx. Similar posture as well as similar body composition probably means similar behavior and diet.
The first thing anyone notices with regard to Spinosaurs like Baryonyx and Subhomimus is the posture. The reason that people always notice the posture is because it is always the same in mounted specimens, casts, and models. Hindlimbs are slightly bent, tail straight out, torso angled downward and forward, with the mouth agape and the forelimbs splayed open, palms of the hands facing one another. The constantly used posture leads to confusion over the type of animal on display. Baryonyx being more widely known than Suchomimus, many Suchomimus skeletons are misnamed and mistaken as Baryonyx. Similar posture as well as similar body composition probably means similar behavior and diet.
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Friendly but Terrible
Even if Suchomimus was a fisherman and would not terrorize humans unless they intruded on its territory, it is a terribly vicious looking dinosaur. Suchomimus does, however, have a pretty good amount of popularity and sway with kids, which tends to be the rule rather than the exception when we are talking about the bigger more frightening looking dinosaurs. This popularity has, thankfully, lead to the creation of kid friendly information pages like the ones at Enchanted Learning and KidsDinos. There are some cartoons, some models/toys, and a multitude of books mention and/or discuss Suchomimus. In terms of the coloring pages I like to share on Sundays, there are a few that are interesting. One that I discovered is rather strange, but it is also the best one; many if not all of the other black and white images are of Spinosaurus, Baryonyx, or are not intended to be coloring pages. The size comparison chart at the Natural History Museum of London fact page could probably be used for coloring also. One other thing that should be noted on this family related day, is that there are many Suchomimus skeleton casts in museums; a trip to the museum and a picnic are great family activities of course. Some museums have a permanent display of Suchomimus while others, such as the Sternberg here in Hays, KS. had one on display while they housed the Giants: African Dinosaurs exhibition that has been traveling between museums since 2011; it was housed in the Sternberg during the summer of 2012 and I missed it due to my moving in and then settling in, total bummer.
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Suchomimus in Action
Suchomimus pops up in a lot of cartoons I have found; or clips anyway. It has also shown up in a couple of older video games. Not many of the cartoon clips get more dramatic than the one included here. That is some violent cartoon action; it is probably still not considered mature or even teen because the violence is between dinosaurs, but it is still pretty violent for a kids' show. Violence kind of drives television though, and I love a good action movie or creature fight (Godzilla was the best movie ever when I was 7), so I am not really complaining. The only drawback to these video games and cartoons, however, is that they are not informative past a certain point. Dinosaur King gives some facts before the fighting is illustrated and sometimes video games also give a small amount of facts, but what we are really lacking here is a good documentary style video. There really is not much in that vein. Enjoy the cartoon fight, however, it is fun to watch.
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Prowling for Beachgoers
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©Walter Myers (www.arcadiastreet.com) |
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Jaws and Families
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©Scott Anselmo, right maxilla |
The hand claws and much of the general shape of the animals are what have tied them closely together. However, Sereno et al (1998) detailed characters that were unique to Suchomimus and separated the two genera. One such character was the number of teeth. Baryonyx possessed 96 teeth in its jaws while Suchomimus houses approximately 130. Our mouths are not designed anything like these giant fish seines. Despite the image of spinosaur mouths as tooth lined nets do not forget that some of the diet of Suchomimus and others in the family tree probably came from carrion as well as fresh caught fish. What is the notch for in the jaw though?
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©Scott Anselmo,mandible |
Reference for the day:
Sereno, P.C.; Beck, A.L.; Dutheil, D.B.; Gado, B.; Larsson, H.C.E.; Lyon, G.H.; Marcot, J.D.; Rauhut, O.W.M.; Sadleir, R.W.; Sidor, C.A.; Varricchio, D.D.; Wilson, G.P; and Wilson, J.A. (1998). "A long-snouted predatory dinosaur from Africa and the evolution of spinosaurids". Science 282 (5392): 1298–1302
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Statues and Electronics
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From the Jurassic Park Wiki, artist uncredited |
Then there are the people that build the Suchomimus, or other dinosaurs, from the ground up as much as is possible without building their own wireframes and code. These are always fun to see and interesting because the tools are set and it takes a lot of ingenuity and ability to accurately use them to create a known dinosaur. I have a healthy respect for the Spore creators that post fairly nice creatures they molded. I picked the one I think is the best out of a very big list, for a dinosaur in Spore, of Suchomimus models:
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You and Your Strange Teeth
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©Nobu Tamura |
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Heterodontosaurus Tusks!
Does sexual dimorphism exist in Heterodontosaurus specimens? The tusks may be a good indication of sexual dimorphism if many more specimens of varying ages (developmentally not geologically) are recovered. Tusks in extant animals, in addition to serving dietary and defensive purposes, are sometimes used to differentiate between sexes. Males with bigger tusks could be seen as older, fitter, or just more handsome in general to the females that are usually impressed by such ornamentations. The tusks of Heterodontosaurus are not enormous though they are a decent and respectable size for a dog sized ornithischian. In a range from warthog to walrus they are much closer to warthog sized teeth. Their position in the skull allows them to have been used in intraspecific rivalries, interspecific fighting, and even to crack open the tougher staples of the Heterodontosaurus diet such as termite mounds.
The skull even appears to have adapted to seating the tusks within the mouth, mostly, as opposed to the way tusks in most extant animals are situated. Picture a walrus or a warthog, even an elephant's tusks and how they protrude from the skull and mouth in a way in which they are never actually concealed by lips, cheeks, or any other fleshy anatomy. The tusks of the Heterodontosaurus were modest enough that they could have been concealed within the cavity of the cheek provided it was stretched that far forward on the skull. Those little tusks even appear to have a position within the junction of the maxilla and premaxilla to accommodate them. Biting down on something seated in that notch with the tusk piercing it from the other side could have caused great trauma, to the limbs of other animals especially, or destruction, in the case of the termite mounds mentioned before. There would have been great risk to the Heterodontosaurus itself, however. Much like any other animal with a tusk it risked breaking the tusk when using it for any activity. Unlike mammals, however, dinosaurs have batteries of teeth, so it is plausible, though perhaps unknown at this time to my knowledge, that the tusks of Heterodontosaurus could have been regrown if broken.
The skull even appears to have adapted to seating the tusks within the mouth, mostly, as opposed to the way tusks in most extant animals are situated. Picture a walrus or a warthog, even an elephant's tusks and how they protrude from the skull and mouth in a way in which they are never actually concealed by lips, cheeks, or any other fleshy anatomy. The tusks of the Heterodontosaurus were modest enough that they could have been concealed within the cavity of the cheek provided it was stretched that far forward on the skull. Those little tusks even appear to have a position within the junction of the maxilla and premaxilla to accommodate them. Biting down on something seated in that notch with the tusk piercing it from the other side could have caused great trauma, to the limbs of other animals especially, or destruction, in the case of the termite mounds mentioned before. There would have been great risk to the Heterodontosaurus itself, however. Much like any other animal with a tusk it risked breaking the tusk when using it for any activity. Unlike mammals, however, dinosaurs have batteries of teeth, so it is plausible, though perhaps unknown at this time to my knowledge, that the tusks of Heterodontosaurus could have been regrown if broken.
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Little Dinosaur for Little Kids
Heterodontosaurus was a little dinosaur. Little kids like little dinosaurs, but that is not all of the appeal. The appeal is also that it is just an interesting little dinosaur and there is a lot of information out there for kids. It is always nice to have a lot of information out there to share on kid friendlyfact pages. Sadly, today, there is not a really good coloring page. However, there is this really neat looking Steve Bledsoe drawing, but I did not ask permission to post, so you only get a link today. Get out there and enjoy Father's Day. Doodle your own Heterodontosaurus with dad and if you like, send it over this way and I will post it. I would like to make a folder (and continuous blog post) for all the drawings kids do, that would be nice.
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Scary Flesh
The fleshed model of Heterodontosaurus that you my see in timelapse images of the construction of the model is a little frightening. The horror of it is no doubt due in part to the rather mammalian appearance of the model at first glance. Actually watching the video takes away from the creepiness of the model a tiny bit, but only because you get to see the feathering and skin details get placed on the model. It should also be made apparent that the model in that video is actually of a different genus of dinosaur related to Heterodontosaurus (Pegomastax africanus is depicted by the model). While a different dinosaur it is closely related to Heterodontosaurus and provides the only fleshed out model that accurately represents the family at the time being; hopefully better models of Heterodontosaurus will be created independently at some point. For that time being, however, enjoy, or at least try to not be freaked out by, this Heterodontosaurid model being created:
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Heterodontosaurus on the Newsstand
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Artist uncredited |
More recently two articles have been published, still a small number considering the research done on other organisms, that detail cranial anatomy as well as morphology. These are good papers and, the paper on juvenile cranial anatomy I can say for certain, because it is a JVP article so I could access it and read it, is quite informative and not a bad read at all. The other paper, focusing on morphology and cranial anatomy is available through Wiley's publication of the Zoological Journal of the Linnaen Society. The article was published in 2011 and was inspired by notes and illustrations of Alan Charig that were passed on to the lead author David Norman after Charig's passing in 1997. That makes it a touching story behind an interesting anatomical discussion I think.
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Everything is Bland
Heterodontosaurus, aside from the tusks and other two types of teeth, is a very typical little dinosaur. Heterodontosaurus was small enough to be compared to a modern retriever and would have been susceptible to attacks from larger dinosaurs but also animals as small as Coelophysis (not that Coelophysis was the smallest predatory dinosaur ever). The little tusks, or fangs as they have been alternately described, would have come into play in predator-prey situations. Undoubtedly the tusks could be used as defensive weaponry against attackers; however, the idea that the tusks represent a characteristic of a sexually dimorphic tradition in Heterodontosaurus means that the females of the species (or males if we reverse the assumption of dimorphism) were much less able to defend themselves in terms of anatomical weaponry.
In terms of where those little tusks/fangs were seated in the skull we are talking about the caudal body of the premaxilla. The bone itself is adjusted so that the teeth sit comfortably into the notch, or diastema, that is formed by this adaptation of the bone. The maxilla itself joins the premaxilla in a way that forms the caudal wall of the diastema where the tusks are situated. The skull is actually, despite the images shared showing the skull earlier in the week, not as exaggerated as we have assumed. Reconstructed in three dimensions, this formation in the skull that houses the tusks is really little more than a narrowing of the solid oral cavity in the area where the teeth are pronounced upward into the mouth. See below:
Reference to check out today:
In terms of where those little tusks/fangs were seated in the skull we are talking about the caudal body of the premaxilla. The bone itself is adjusted so that the teeth sit comfortably into the notch, or diastema, that is formed by this adaptation of the bone. The maxilla itself joins the premaxilla in a way that forms the caudal wall of the diastema where the tusks are situated. The skull is actually, despite the images shared showing the skull earlier in the week, not as exaggerated as we have assumed. Reconstructed in three dimensions, this formation in the skull that houses the tusks is really little more than a narrowing of the solid oral cavity in the area where the teeth are pronounced upward into the mouth. See below:
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From The Dinosauria |
Weishampel, D. B. and Witmer, L. M. (1990). Heterodontosauridae. in Weishampel, D. B., Dodson, P., & Osmólska, H. (Eds.). Pgs. 486-497. The Dinosauria. Univ of California Press.
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Vaguely Popular
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Kind of a big Iguana until you get to the head |
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Running Through the Delta
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©Nobu Tamura |
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Speculative Craniums
The cranial elements of Deltadromeus are missing entirely. It is certainly not rare for skeletal elements of dinosaurs to be missing and skulls are high on the list of frequently AWOL skeletal elements. An interesting question of taphonomy can be posed when we consider this. What happens to all of those heads that are missing from the fossil record? Are heads washed away easily when the flesh is gone or are they dragged away by scavengers that do not wish to share the brains ("zombie" dinosaur scavengers)? Did heads get buried less frequently than other elements of the body? Whatever the reason, it has led to the recovery of Deltadromeus specimens made up entirely, to this point, of post-cranial material. Thus, all of the representations of the head of Deltadromeus are speculations based on presumably related taxa. Typically this means that the skulls are based on other ceratosaurs. This has led to a variety of ornamentation on the skulls of illustrated and mounted Deltadromeus. Typically a ridge of material has been placed above each orbit, similar to those seen in Ceratosaurus.
The cranial ornaments change over time. They are basically all very similar, but the positions, angles, and overall shape of the crests above the orbits change over time and between illustrators and model makers. This reflects not only personal preferences but changing hypotheses concerning the exact nature and potential construction of the skull of Deltadromeus. It is still not certain that Deltadromeus is a ceratosaur also and the skull may therefore have absolutely no ornamentation at all. It is even conceivable that Deltadromeus, as a very basal ceratosaur may lack crests on the skull entirely for that reason. Regardless, there are certain elements of the body of Deltadromeus that are known fairly well. These include the legs and the majority of the caudal vertebrae. The legs and caudal vertebrae are similar to those found in ceratosaurs, thereby aiding the justification that Deltadromeus is a ceratosaur. Seeing as how that is the best evidence and the best fit for known remains, that is where it is positioned in the dinosaur family tree, for now; the legs and may not be telling all there is to know about this interesting dinosaur.
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©Mineo Shiraishi |
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Running for Children
I suppose it is more likely that Deltadromeus would run after children rather than for children. If dinosaurs were around like in Dinotopia cancer runs would raise ridiculous amounts of money I bet though; I like to think dinosaurs had more stamina than the average person. Anyhow, Deltadromeus is a fairly kid friendly dinosaur. There are a number of child oriented fact pages online; About and Age of Dinosaurs have the best ones for today. Kids that cannot put down electronics can play some Dinosaur King with Deltadromeus or they can pick up the card game and play with Deltadromeus in a slightly less animated manner. Children that would rather use all of their imagination instead of letting the electronics do some of it for them could grab a Deltadromeus model/toy crafted by Safari LTD; though it is designed for the Carnegie collection which means that its up a little higher in the price world. The Safari LTD model is skull ornamentation free, another interpretation of what the skull may look like should we ever find one. If your young ones would rather read there is a book called Deltadromeus and other Shoreline Dinosaurs out there that they could pick up and read through. There are a lot of resources for kids to enjoy for Deltadromeus.
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